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(BA Anthropology 1996, Pomona College ; MA Anthropology, 2011 ³Ô¹ÏÍø of Colorado, PhD Anthropology, 2019 ³Ô¹ÏÍø of Colorado). Paine is interested in the dispersal and speciation patterns of the australopithecines, focusing on the broad ecological and biogeographical context in which they occurred. His advisor was Matt Sponheimer.